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MUDs, MOOs, MUCKs and MUSHes.
Usenet in general
sodaconstructor.com
memepool
ectoplasmosis
Boing Boing when it was great
An early virtual world called WorldsAway, technically still exists at VZones but I don't think it's possible to make an account now
Hotwired
Suck.com and early discussion side adjunct Plastic.com
TimmyBigHands, short-lived humor magazine from the Mystery Science Theater 3000 people shortly after the show folded the second time.
The Sci-Fi Channel's website and MTV.com, both started with the hopes of becoming a substantial part of the World Wide Web by getting in early, then shuttered when owners lost interest or believed the lies that social media was where the only thing that mattered any more. Also Cartoon Network's website, which was once a joy. Adult Swim's website hangs on, but is a shadow of its former self, and doesn't host forums any longer.
The Usenet MSTing archives at pinky.wtower.com
I mean, they're still out there. Not all of them, and the player populations aren't as large as in their heyday.
https://www.mudverse.com/
Was just playing a mud earlier today!
I know, but it's not nearly the same kind of thing as back when they were big, sadly.
One thing that still exists is WWWF Grudge Match, a likely inspiration for MTV's Celebrity Deathmatch, a crowdsourced humor site where they put different pop culture figures against each other in textual battles and people contributed their takes and votes on who would win. It hasn't updated in over two decades though. It even spawned a book!
(I am slightly responsible for its continued existence. I noticed that it had gone offline a while back. I happened to have Chris, one of its founders, in my contacts list, so I sent him a message about it. Turns out to have been an expired credit card, a quick change of backing funds and GRUDGE MATCH LIVES AGAIN!)
Also, The Conservatron, another crowdsource humor site, home of the Evil Black Marble from Marble Madness.
It seems like the Internet Oracle has finally, at last, succumbed to linkrot.
Two sites that still exist are Everything, H2G2 and the homepage of venerable roguelike game Nethack!